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Gill, BA, S Kurukura, AA Hassan, JR Goheen, WJ Kress, M Kuzmina, RM Pringle, and TR Kartzinel. 2018. Plant DNA-barcode library and community phylogeny for a semi-arid East African savanna. Molecular Ecology Resources: in press.

Pansu, J, JA Guyton, AB Potter, JL Atkins, JH Daskin, B Wursten, TR Kartzinel, and RM Pringle. 2018. Trophic ecology of large herbivores in a reassembling African ecosystem. Journal of Ecology: in press.

Reese, AT, FC Pereira, A Schintlmeister, D Berry, M Wagner, LP Hale, HK Durand, X Zhou, R Premont, AM Diehl, TM O’Connell, SC Alberts, TR Kartzinel, RM Pringle, RR Dunn, JP Wright, and LA David. Microbial nitrogen limitation in the mammalian large intestine. Nature Microbiology 3:1441-1450. [link].

Daskin, JH and RM Pringle. 2018. Warfare and wildlife declines in Africa’s protected areas. Nature 553:328-332. [PDF]

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Pringle, RM. 2018. Ecology: megaherbivores homogenize the landscape of fear. Current Biology 28:R835-R837.

Louthan, AM, RM Pringle, JR Goheen, TM Palmer, WF Morris, and DF Doak. 2018. Aridity weakens population-level effects of multiple species interactions in Hibiscus meyeri. Proc. Natl. Acad. Sci. USA 115:543-548. [PDF]

Coverdale, TC, JR Goheen, TM Palmer, and RM Pringle. 2018. Good neighbors make good defenses: associational refuges reduce defense investment in African savanna plants. Ecology 99:1724-1736. [PDF]

Titcomb, G, RM Pringle TM Palmer, HS Young. 2018. What explains tick proliferation following large-herbivore exclusion? Proceedings of the Royal Society B 286:20180612.

Goheen, JR, DJ Augustine, KE Veblen, DM Kimuyu, TM Palmer, LM Porensky, RM Pringle, J Ratnam, C Riginos, M Sankaran, AT Ford, AA Hassan, R Jakopak, TR Kartzinel, S Kurukura, AM Louthan, WO Odadi, TO Otieno, AM Wambua, HS Young, and TP Young. 2018. Conservation lessons from large-mammal manipulations in East African savannas: the KLEE, UHURU, and GLADE experiments. Annals of the New York Academy of Sciences 1429:31-49. [PDF]


Pringle, RM. 2017. Upgrading protected areas to conserve wild biodiversity. Nature 546:91-99. [PDF] [em Português] [en Espanñol]

Pringle, RM. 2017. How large herbivores subsidize aquatic food webs in African savannas. Proc. Natl. Acad. Sci. USA 114:7489-7491. [PDF]

Pringle, RM and CE Tarnita. 2017. Spatial self-organization of ecosystems: integrating multiple mechanisms of regular-pattern formation. Annual Review of Entomology 62:359-377. [PDF]

Tarnita, CE, JA Bonachela, E Sheffer, JA Guyton, TC Coverdale, RA Long, and RM Pringle. 2017. A theoretical foundation for multi-scale regular vegetation patterns. Nature 541:398-401. [PDF]

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Long, RA, A Wambua, JR Goheen, TM Palmer, and RM Pringle. 2017. Climatic variation modulates the indirect effects of large herbivores on small-mammal habitat use. Journal of Animal Ecology 86:739-748. [PDF]

Palmer, TM, C Riginos, R Damiani, N Morgan, J Lemboi, J Lengingiro, J Ruiz Guajardo, and RM Pringle. 2017. Influence of neighboring plants on the dynamics of an ant-acacia protection mutualism. Ecology: doi:10.1002/ecy.2008.

Titcomb, G, BF Allan, T Ainsworth, L Henson, T Hedlund, RM Pringle, TM Palmer, L Njoroge, MG Campaña, R Fleischer, JN Mantas, and HS Young. 2017. Interacting effects of wildlife loss and climate on ticks and tick-borne disease. Proceedings of the Royal Society B: 286:20170475.

Young, HS, DJ McCauley, R Dirzo, CL Nunn, B Agwanda, M Campana, R Fleischer, EO Castillo, D Salked, E Lambin, JR Goheen, TM Palmer, RM Pringle, TP Young, and KM Helgen. 2017. Interacting effects of land-use and climate on rodent-borne disease in central Kenya. Philosophical Transactions of the Royal Society B 372:20160116. [PDF]

Pellegrini, AFA, RM Pringle, N Govender, and LO Hedin. 2017. Woody plant biomass and carbon exchange depend on elephant-fire interactions in African savanna. Journal of Ecology 105:111-121. [PDF]

Ripple, WJ, and 42 co-authors. 2017. Conserving the world’s megafauna and biodiversity: the fierce urgency of now. BioScience 67:197-200. [PDF]


Pringle, RM, KM Prior, TM Palmer, TP Young, and JR Goheen. 2016. Large herbivores promote habitat specialization and beta diversity of African savanna trees. Ecology 97:2640-2657. [PDF]

Coverdale, TC, TR Kartzinel, KL Grabowski, RK Shriver, AA Hassan, JR Goheen, TM Palmer, and RM Pringle. 2016. Elephants in the understory: opposing direct and indirect effects of consumption and ecosystem engineering by megaherbivores. Ecology 97:3219-3230. [PDF]

Ripple, WJ, and 42 co-authors. Saving the world’s terrestrial megafauna. BioScience 66:807-812. [PDF]

Daskin, JH and RM Pringle. 2016. Does primary productivity modulate the indirect effects of large herbivores? A global meta-analysis. Journal of Animal Ecology 85:857–868. [PDF]

Daskin, JH, M Stalmans, and RM Pringle. 2016. Ecological legacies of civil war: 35-year increase in savanna tree cover following wholesale large-mammal declines. Journal of Ecology 104:79-89 [PDF]



Pringle, RM, DM Kimuyu, RL Sensenig, TM Palmer, C Riginos, KE Veblen, TP Young. 2015. Synergistic effects of fire and elephants on arboreal animals in an African savanna. Journal of Animal Ecology 84:1637-1645. [PDF]

Kartzinel, TR, PA Chen, TC Coverdale, DL Erickson, WJ Kress, ML Kuzmina, DI Rubenstein, W Wang, and RM Pringle. 2015. DNA metabarcoding illuminates dietary niche partitioning by African large herbivores. Proc. Natl. Acad. Sci. USA 112:8019-8024. [PDF] [PNAS Commentary by Dave Tilman & Elizabeth Borer]


Bonachela, JA, RM Pringle, E Sheffer, TC Coverdale, JA Guyton, KK Caylor, SA Levin, CE Tarnita. 2015. Termite mounds can increase the robustness of dry land ecosystems to climatic change. Science 347:651-655. [PDF]

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Fraser, LH, and 58 co-authors. 2015. Worldwide evidence supporting a unimodal relationship between productivity and plant species richness. Science 349:302-305. [PDF]

Kartzinel, TR and RM Pringle. 2015. Molecular detection of invertebrate prey in vertebrate diets: trophic ecology of Caribbean island lizards. Molecular Ecology Resources 15:903-914. [PDF]

Ford, AT, JR Goheen, DJ Augustine, MF Kinnaird, TG O’Brien, TM Palmer, RM Pringle, and R Woodroffe. 2015. Recovery of African wild dogs suppresses prey but does not trigger a trophic cascade. Ecology 96:2705-2714. [PDF]

Ceballos, G, PR Ehrlich, AD Barnosky, A Garcia, RM Pringle, and TM Palmer. 2015. Accelerated modern human-induced species losses: entering the sixth mass extinction. Science Advances. 1:e1400253. [PDF]

Young, HS, DJ McCauley, R Dirzo, JR Goheen, B Agwanda, AW Ferguson, SN Kinyua, MM McDonough, TM Palmer, RM Pringle, TP Young, and KM Helgen. 2015. Context-dependent effects of large-wildlife declines on small mammal communities in central Kenya. Ecological Applications 25:348-360. [PDF]

Pringle, RM. 2015. Review of “How to do ecology: a concise handbook (2nd edition)” by R Karban, M Huntzinger, & IS Pearse. Quarterly Review of Biology 90:324-325. [PDF]


Ford, AT, JR Goheen, TO Otieno, L Bidner, L Isbell, TM Palmer, R Woodroffe, D Ward, and RM Pringle. 2014. Large carnivores make savanna tree communities less thorny. Science 346:346-349. [PDF]

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Tarnita, CE, TM Palmer, and RM Pringle. Colonization and competition dynamics can explain incomplete sterilization parasitism in ant-plant symbioses. Ecology Letters 17:1290-1298. [PDF]

Pringle, RM, JR Goheen, TM Palmer, GK Charles, E DeFranco, R Hohbein, AT Ford, B Torto, and CE Tarnita. Low functional redundancy among sympatric large herbivores in regulating an encroaching shrub (Solanum campylacanthum) in African savannah. Proceedings of the Royal Society B 281:20140390. [PDF]

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Louthan, AM, DF Doak, JR Goheen, TM Palmer, and RM Pringle. 2014. Mechanisms of plant-plant interactions: concealment from herbivores is more important than abiotic-stress mediation in an African savanna. Proceedings of the Royal Society B 281:20132647. [PDF]

Ngatia, LW, KR Reddy, PKR Nair, RM Pringle, TM Palmer, and BL Turner. 2014. Seasonal patterns in decomposition and nutrient release from East African savanna grasses grown under contrasting nutrient conditions. Agriculture, Ecosystems & Environment 188:12-19. [PDF]

Mendenhall, CD, G Ceballos, LO Frishkoff, J Pacheco, G Santos-Barrera, PR Ehrlich, GC Daily, and RM Pringle. 2014. Countryside biogeography of Neotropical reptiles and amphibians. Ecology 95:856-870. [PDF]

Kartzinel, TR, JR Goheen, GK Charles, E DeFranco, JE Maclean, TO Otieno, TM Palmer, and RM Pringle. 2014. Plant and small-mammal responses to large-herbivore exclusion in an African savanna: the first five years of the UHURU experiment. Ecology 95:787. [PDF]

Pringle, RM. 2014. Where the wild beasts are. Review of “Serengeti story: life and science in the world’s greatest wildlife region” by ARE Sinclair. Ecology 95:583-584. [PDF]


Palmer TM, ML Stanton, TP Young, JR Goheen, JS Lemboi, and RM Pringle. 2013. A role for indirect facilitation in maintaining diversity in a guild of African acacia ants. Ecology 94:1531-1539. [PDF]

Donihue, CM, LM Porensky, J Foufopoulos, C Riginos, and RM Pringle. 2013. Glade cascades: indirect legacy effects of pastoralism enhance the abundance and spatial structuring of arboreal fauna. Ecology 94:827-837. [PDF]

Louthan, AM, DF Doak, JR Goheen, TM Palmer, and RM Pringle. 2013. Climatic stress mediates the impacts of herbivory on plant population structure and components of individual fitness. Journal of Ecology 101:1074-1083. [PDF]

Young, HS, DJ McCauley, KM Helgen, E Otarola-Castillo, JR Goheen, TM Palmer, RM Pringle, TP Young, and R Dirzo. 2013. Effects of mammalian herbivore declines on plant communities: observations and experiments in an African savanna. Journal of Ecology 101:1030-1041. [PDF]

Goheen, JR, TM Palmer, GK Charles, KM Helgen, SN Kinyua, JE Maclean, HS Young, and RM Pringle. 2013. Piecewise disassembly of a large-herbivore community across a rainfall gradient: the UHURU experiment. PLOS One 8:e55192. doi:10.1371/journal.pone.0055192 [PDF]


Pringle, RM. 2012. How to be manipulative. American Scientist 100:30-37. [PDF]


Losos, JB and RM Pringle. 2011. Competition, predation, and natural selection in island lizards. Nature 475:E1-2. [PDF]

Pringle, RM, TM Palmer, JR Goheen, DJ McCauley, F. Keesing. 2011. Ecological importance of large herbivores in the Ewaso ecosystem. Pp. 43-53 in NJ Georgiadis (ed), Conserving Wildlife in African Landscapes: Kenya’s Ewaso Ecosystem. Smithsonian Contributions to Zoology No. 632. Smithsonian Institution Scholarly Press, Washington DC. [PDF]

Pringle, RM. 2011. Nile perch. Pp. 484-488 in D. Simberloff and M. Rejmánek (eds), Encyclopedia of Biological Invasions. University of California Press, Berkeley, CA. [PDF]


Palmer, TM, DF Doak, ML Stanton, JL Bronstein, ET Kiers, TP Young, JR Goheen, and RM Pringle. 2010. Synergy of multiple partners, including freeloaders, increases host fitness in a multispecies mutualism. Proc. Natl. Acad. Sci. USA 107:17234-17239. [PDF]

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Pringle, RM, DF Doak, AK Brody, R Jocqué, and TM Palmer. 2010. Spatial pattern enhances ecosystem functioning in an African savanna. PLOS Biology 8:e1000377. doi:10.1371/journal.pbio.1000377 [PDF] [PLOS Biology Commentary by Oswald J Schmitz]

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Pringle, RM. 2010. The costs of large-mammal extinctions. Pp. 52-53 in NS Sodhi and PR Ehrlich (eds), Conservation Biology for All. Oxford University Press, Oxford, UK. [PDF]

Pringle, RM. 2010. Review of “Conservation Psychology: Understanding and Promoting Human Care for Nature” by S Clayton and G Myers. Environmental Conservation 37:222. [PDF]


Pringle, RM, M Syfert, JK Webb, and R Shine. 2009. Quantifying historical changes in habitat availability for endangered species: use of pixel- and object-based remote sensing. Journal of Applied Ecology 46:544-553. [PDF]

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Angilletta, MJ, MW Sears, and RM Pringle. 2009. The spatial dynamics of nesting behavior: lizards shift microhabitats to construct nests with beneficial thermal properties. Ecology 90:2933-2939. [PDF]

Roman, J, PR Ehrlich, RM Pringle, and JC Avise. 2009. Facing extinction: nine steps to save biodiversity. Solutions 1:27-37. [PDF]

Webb, JK, RM Pringle, and R Shine. 2009. Intraguild predation, thermoregulation, and microhabitat selection by snakes. Behavioral Ecology 20:271-277. [PDF]

Pringle, RM. 2009. Review of “Paradise Found: Nature in America at the Time of Discovery,” by S Nicholls. Environmental Conservation 36:173-174. [PDF]


Pringle, RM and K Fox-Dobbs. 2008. Coupling of canopy and understory food webs by ground-dwelling predators. Ecology Letters 11:1328-1337. [PDF]

Ehrlich, PR and RM Pringle. 2008. Where does biodiversity go from here? A grim business-as-usual forecast and a hopeful portfolio of partial solutions. Proc. Natl. Acad Sci. USA 105:11579-11586. [PDF]

Pejchar, L., RM Pringle, J Ranganathan, JR Zook, G Duran, and GC Daily. 2008. Birds as agents of seed dispersal in a human-dominated landscape in Costa Rica. Biological Conservation 141:536-544. [PDF]

Palmer, TM, ML Stanton, TP Young, JR Goheen, RM Pringle, and R Karban. 2008. Putting ant-acacia mutualisms to the fire. Science 319:1760-1761. [PDF]

Pringle, RM. 2008. Elephants as agents of habitat creation for small vertebrates at the patch scale. Ecology 89:26-33. [PDF]


Palmer, TM, ML Stanton, TP Young, JR Goheen, RM Pringle, and R Karban. 2008. Breakdown of an ant-plant mutualism follows the loss of large herbivores from an African savanna. Science 319:192-195. [PDF]

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Pringle, RM, TP Young, DI Rubenstein, and DJ McCauley. 2007. Herbivore-initiated interaction cascades and their modulation by productivity in an African savanna. Proc. Natl. Acad. Sci. USA 104:193-197. [PDF] [PNAS Commentary by Andrew Larson & Bob Paine]

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Chan, KMA, RM Pringle, J Ranganathan, CL Boggs, YE Chan, PR Ehrlich, PK Haff, NE Heller, K Al-Khafaji, and D. MacMynowski. 2007. When agendas collide: human welfare and biological conservation. Conservation Biology 21:59-68. [PDF]


McCauley, DJ, F Keesing, TP Young, BF Allan, and RM Pringle. 2006. Indirect effects of large herbivores on snakes in an African savanna. Ecology 87:2657-2663. [PDF]

Pringle, RM. 2006. Thinking anthropologically about nature. Review of “People and Nature,” by EF Moran. Trends in Ecology & Evolution 21:483-484. [PDF]


Pringle, RM. 2005. The Nile perch in Lake Victoria: local responses and adaptations. Africa 75:510-538. [PDF]

Pringle, RM. 2005. The origins of the Nile perch in Lake Victoria. BioScience 55:780-787. [PDF]

Webb, JK, R Shine, and RM Pringle. 2005. Canopy removal restores habitat quality for an endangered snake in a fire-suppressed landscape. Copeia 2005:894-900. [PDF]


Webb, JK, RM Pringle, and R Shine. 2004. How do nocturnal snakes select diurnal retreat sites? Copeia 2004: 919-925. [PDF]


Pringle, RM, JK Webb, and R Shine. 2003. Canopy structure, microclimate, and habitat selection by the nocturnal snake Hoplocephalus bungaroides. Ecology 84:2668-2679. [PDF]